GRNA / GreenLight Biosciences Inc - SEC-Einreichungen, Jahresbericht, Proxy Statement

GreenLight Biosciences Inc
US ˙ NASDAQ
DIESES SYMBOL IST NICHT MEHR AKTIV

Basisstatistiken
CIK 1822691
SEC Filings
All companies that sell securities in the United States must register with the Securities and Exchange Commission (SEC) and file reports on a regular basis. These reports include company annual reports (10K, 10Q), news updates (8K), investor presentations (found in 8Ks), insider trades (form 4), ownership reports (13D, and 13G), and reports related to the specific securities sold, such as registration statements and prospectus. This page shows recent SEC filings related to GreenLight Biosciences Inc
SEC Filings (Chronological Order)
Auf dieser Seite finden Sie eine vollständige, chronologische Liste der SEC-Einreichungen, mit Ausnahme der Eigentumseinreichungen, die wir an anderer Stelle bereitstellen.
August 11, 2023 15-12G

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 15 CERTIFICATION AND NOTICE OF TERMINATION OF REGISTRATION UNDER SECTION 12(g) OF THE SECURITIES EXCHANGE ACT OF 1934 OR SUSPENSION OF DUTY TO FILE REPORTS UNDER SECTIONS 13

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 15 CERTIFICATION AND NOTICE OF TERMINATION OF REGISTRATION UNDER SECTION 12(g) OF THE SECURITIES EXCHANGE ACT OF 1934 OR SUSPENSION OF DUTY TO FILE REPORTS UNDER SECTIONS 13 AND 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934. Commission File Number 001-39894 GreenLight Biosciences Holdings, PBC (Exact name of registrant as

July 28, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Insud Pharma S.L. - AMENDMENT TO SCHEDULE 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. 1)* Greenlight Biosciences Holdings, PBC (Name of Issuer) Common stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) c/o Ana Mondedeu Insunza Insud Pharma, S.L. Calle de Manuel Pombo Angulo, No. 28, 3rd Floor 28050 Mad

July 28, 2023 EX-3.2

BYLAWS GREENLIGHT BIOSCIENCES HOLDINGS, PBC., a public benefit corporation ARTICLE I.

Exhibit 3.2 BYLAWS OF GREENLIGHT BIOSCIENCES HOLDINGS, PBC., a public benefit corporation ARTICLE I. OFFICES Section 1. Registered Office. The registered office shall be at the office of The Corporation Trust Company, 1209 Orange Street, Wilmington, Delaware 19801. Section 2. Other Offices. The corporation may also have offices at such other places both within and without the State of Delaware as

July 28, 2023 8-K/A

Changes in Control of Registrant, Departure of Directors or Certain Officers; Election of Directors; Appointment of Certain Officers; Compensatory Arrangements of Certain Officers, Amendments to Articles of Incorporation or Bylaws; Change in Fiscal Year, Material Modification to Rights of Security Holders, Notice of Delisting or Failure to Satisfy a Continued Listing Rule or Standard; Transfer of Listing, Financial Statements and Exhibits, Completion of Acquisition or Disposition of Assets, Other Events

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K/A (Amendment No. 1) CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): July 24, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdictio

July 28, 2023 EX-99.2

GreenLight Announces Completion of Merger with Fall Line Endurance Fund

Exhibit 99.2 GreenLight Announces Completion of Merger with Fall Line Endurance Fund Lexington, Mass., July 24, 2023 – GreenLight Biosciences Holdings, PBC (NASDAQ: GRNA) (“GreenLight” or the “Company”) today announced the closing of its previously announced merger with SW MergerCo, Inc. (“Merger Sub”) and SW ParentCo, Inc. (“Parent”), pursuant to which Merger Sub merged with and into GreenLight w

July 28, 2023 EX-99.1

GreenLight Biosciences Announces Expiration of Tender Offer Period for Outstanding Shares Completion of Previously Announced Merger and Going Private Transaction Expected July 20, 2023

Exhibit 99.1 GreenLight Biosciences Announces Expiration of Tender Offer Period for Outstanding Shares Completion of Previously Announced Merger and Going Private Transaction Expected July 20, 2023 Lexington, Mass. – July 20, 2023 – GreenLight Biosciences Holdings, PBC (NASDAQ: GRNA) (“GreenLight” or the “Company”) today announced that it is initiating the final steps in closing its previously ann

July 28, 2023 EX-3.1

THIRD AMENDED AND RESTATED CERTIFICATE OF INCORPORATION GREENLIGHT BIOSCIENCES HOLDINGS, PBC A PUBLIC BENEFIT CORPORATION ARTICLE 1

Exhibit 3.1 THIRD AMENDED AND RESTATED CERTIFICATE OF INCORPORATION OF GREENLIGHT BIOSCIENCES HOLDINGS, PBC A PUBLIC BENEFIT CORPORATION ARTICLE 1 The name of the public benefit corporation is GreenLight Biosciences Holdings, PBC. ARTICLE 2 The address of the Corporation’s registered office in the State of Delaware is c/o The Corporation Trust Company, 1209 Orange Street, Wilmington, Delaware 1980

July 28, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Boscolo Intervest Ltd - SC 13D/A Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. 1) GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Rafael Urquia II, Esq. c/o Fox Horan & Camerini LLP 885 Third Avenue, 17th Floor New York, NY 10022 (2

July 27, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / KODIAK VENTURE PARTNERS III LP - SC 13D/A Activist Investment

SC 13D/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D/A Under the Securities Exchange Act of 1934 (Amendment No. 2)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) David Furneaux Kodiak Ventures Management Company, Inc. 11 Peter Grover Rd Bethel, ME 0421

July 27, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / S2G Advisors, LLC - SC 13D/A Activist Investment

SC 13D/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. 1) GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Lisa J. Forbes 110 N.W. 2nd Street, Suite 300 Bentonville, AR 72172 (479) 464-1500 (Name, Add

July 26, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / ALEXANDRIA REAL ESTATE EQUITIES, INC. - SCHEDULE 13D/A Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. 1)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Dean A. Shigenaga Chief Financial Officer Alexandria Venture Investments, LLC 26 North Euclid Avenue

July 26, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Jeremy & Hannelore Grantham Environmental Trust - SC 13D/A Activist Investment

SC 13D/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. 1)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Ramsay Ravenel The Jeremy and Hannelore Grantham Environmental Trust 53 State Street, 33rd F

July 26, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / MLS Capital Fund II LP - SC 13D/A Activist Investment

SC 13D/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D/A Under the Securities Exchange Act of 1934 (Amendment No. 1)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Gregory Young MLSCF Management (Labuan), LLP Suite 26.03, Level 26, GTower 199, Jalan Tun

July 26, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / CONTINENTAL GRAIN CO - AMENDMENT NO. 1 TO SCHEDULE 13D Activist Investment

SC 13D/A 1 ea182381-13da1contingreen.htm AMENDMENT NO. 1 TO SCHEDULE 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 SCHEDULE 13D/A (Rule 13d-101) INFORMATION TO BE INCLUDED IN STATEMENTS FILED PURSUANT TO § 240.13d-l(a) AND AMENDMENTS THERETO FILED PURSUANT TO § 240.13d-2(a) (Amendment No. 1)* Greenlight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par valu

July 26, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Board of Trustees of The Leland Stanford Junior University - SC 13D/A Activist Investment

SC 13D/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. 1)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Michael Taylor The Board of Trustees of the Leland Stanford Junior University 635 Knight Way

July 25, 2023 EX-99.1

JOINT FILING AGREEMENT

EX-99.1 2 d457540dex991.htm EX-99.1 CUSIP No. 39536G105 EXHIBIT 99.1 JOINT FILING AGREEMENT The undersigned, being duly authorized thereunder, hereby execute this agreement as an exhibit to this Schedule 13D amendment to evidence the agreement of the below-named parties, in accordance with the rules promulgated pursuant to the Securities Exchange Act of 1934, to file this Schedule 13D amendment (i

July 25, 2023 EX-99.3

JOINT FILING AGREEMENT

Exhibit 99.3 JOINT FILING AGREEMENT In accordance with Rule 13d-1(k)(1) promulgated under the Securities Exchange Act of 1934, as amended, the undersigned hereby agree that they are jointly filing this statement on Schedule 13D. Each of them is responsible for the timely filing of such statement and any amendments thereto, and for the completeness and accuracy of the information concerning such pe

July 25, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / RPB VENTURES LLC Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Ruth Eloise Beneby Shirley & Charlotte Streets, P.O. Box N-1175 Bahamas Financial Centre, 2nd Floor Nassau, Bahamas +1 (

July 25, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Morningside Venture Investments Ltd - SC 13D/A Activist Investment

SC 13D/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. 2)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G105 (CUSIP Number) Morningside Venture Investments Limited C/O THC Management Services S.A.M., 2nd Floor, Le Pri

July 24, 2023 EX-3.1

THIRD AMENDED AND RESTATED CERTIFICATE OF INCORPORATION GREENLIGHT BIOSCIENCES HOLDINGS, PBC A PUBLIC BENEFIT CORPORATION ARTICLE 1

EX-3.1 Exhibit 3.1 THIRD AMENDED AND RESTATED CERTIFICATE OF INCORPORATION OF GREENLIGHT BIOSCIENCES HOLDINGS, PBC A PUBLIC BENEFIT CORPORATION ARTICLE 1 The name of the public benefit corporation is GreenLight Biosciences Holdings, PBC. ARTICLE 2 The address of the Corporation’s registered office in the State of Delaware is c/o The Corporation Trust Company, 1209 Orange Street, Wilmington, Delawa

July 24, 2023 S-8 POS

As filed with the Securities and Exchange Commission on July 24, 2023

S-8 POS As filed with the Securities and Exchange Commission on July 24, 2023 Registration No.

July 24, 2023 POS AM

As filed with the Securities and Exchange Commission on July 24, 2023

POS AM As filed with the Securities and Exchange Commission on July 24, 2023 Registration No.

July 24, 2023 EX-99.2

GreenLight Announces Completion of Merger with Fall Line Endurance Fund

EX-99.2 Exhibit 99.2 GreenLight Announces Completion of Merger with Fall Line Endurance Fund Lexington, Mass., July 24, 2023 – GreenLight Biosciences Holdings, PBC (NASDAQ: GRNA) (“GreenLight” or the “Company”) today announced the closing of its previously announced merger with SW MergerCo, Inc. (“Merger Sub”) and SW ParentCo, Inc. (“Parent”), pursuant to which Merger Sub merged with and into Gree

July 24, 2023 EX-3.2

BYLAWS GREENLIGHT BIOSCIENCES HOLDINGS, PBC., a public benefit corporation ARTICLE I.

EX-3.2 Exhibit 3.2 BYLAWS OF GREENLIGHT BIOSCIENCES HOLDINGS, PBC., a public benefit corporation ARTICLE I. OFFICES Section 1. Registered Office. The registered office shall be at the office of The Corporation Trust Company, 1209 Orange Street, Wilmington, Delaware 19801. Section 2. Other Offices. The corporation may also have offices at such other places both within and without the State of Delaw

July 24, 2023 8-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): July 24, 2023 GreenLight Bioscien

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): July 24, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporation)

July 24, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Zahir Fares - SC 13D/A Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. 1)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G105 (CUSIP Number) Fares Zahir c/o TMF Nominees Ltd, 2nd Floor, The Grand Pavilion, Commercial Centre, 802 West Bay Road,

July 24, 2023 EX-99.1

GreenLight Biosciences Announces Expiration of Tender Offer Period for Outstanding Shares Completion of Previously Announced Merger and Going Private Transaction Expected July 20, 2023

EX-99.1 Exhibit 99.1 GreenLight Biosciences Announces Expiration of Tender Offer Period for Outstanding Shares Completion of Previously Announced Merger and Going Private Transaction Expected July 20, 2023 Lexington, Mass. – July 20, 2023 – GreenLight Biosciences Holdings, PBC (NASDAQ: GRNA) (“GreenLight” or the “Company”) today announced that it is initiating the final steps in closing its previo

July 24, 2023 POS AM

As filed with the Securities and Exchange Commission on July 24, 2023

As filed with the Securities and Exchange Commission on July 24, 2023 Registration No.

July 20, 2023 SC 14D9/A

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 14D-9 SOLICITATION/RECOMMENDATION STATEMENT UNDER SECTION 14(d)(4) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 4) GreenLight Biosciences Holdings, PBC (Name of

SC 14D9/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 14D-9 SOLICITATION/RECOMMENDATION STATEMENT UNDER SECTION 14(d)(4) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 4) GreenLight Biosciences Holdings, PBC (Name of Subject Company) GreenLight Biosciences Holdings, PBC (Name of Person(s) Filing Statement) Common Stock, par value $0.0001 per share (Title

July 20, 2023 EX-99.(A)(5)(1)

GreenLight Biosciences Announces Expiration of Tender Offer Period for Outstanding Shares Completion of Previously Announced Merger and Going Private Transaction Expected July 20, 2023

EX-99.(a)(5)(1) Exhibit (a)(5)(1) GreenLight Biosciences Announces Expiration of Tender Offer Period for Outstanding Shares Completion of Previously Announced Merger and Going Private Transaction Expected July 20, 2023 Lexington, Mass. – July 20, 2023 – GreenLight Biosciences Holdings, PBC (NASDAQ: GRNA) (“GreenLight” or the “Company”) today announced that it is initiating the final steps in closi

July 20, 2023 EX-99.(A)(5)(C)

GreenLight Biosciences Announces Expiration of Tender Offer Period for Outstanding Shares Completion of Previously Announced Merger and Going Private Transaction Expected July 20, 2023

EX-99.(a)(5)(C) Exhibit (a) (5) (C) GreenLight Biosciences Announces Expiration of Tender Offer Period for Outstanding Shares Completion of Previously Announced Merger and Going Private Transaction Expected July 20, 2023 Lexington, Mass. – July 20, 2023 – GreenLight Biosciences Holdings, PBC (NASDAQ: GRNA) (“GreenLight” or the “Company”) today announced that it is initiating the final steps in clo

July 20, 2023 SC TO-T/A

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE TO TENDER OFFER STATEMENT UNDER SECTION 14(d)(1) OR 13(e)(1) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 2) GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Name of Subje

SC TO-T/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE TO TENDER OFFER STATEMENT UNDER SECTION 14(d)(1) OR 13(e)(1) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 2) GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Name of Subject Company (Issuer)) SW MergerCo, Inc. a wholly-owned subsidiary of (Name of Filing Person — Offeror) SW ParentCo, Inc. an affiliate of (Nam

July 20, 2023 SC 13E3/A

SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13E-3 (Rule 13e-100) (Amendment No. 4) Rule 13e-3 Transaction Statement Under Section 13(e) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of the Is

SC 13E3/A SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13E-3 (Rule 13e-100) (Amendment No. 4) Rule 13e-3 Transaction Statement Under Section 13(e) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of the Issuer) Fall Line Endurance Fund, LP (and the affiliated entities and other persons listed on the following page) (Name of Persons Filing Stat

July 13, 2023 SC 13E3/A

SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13E-3 (Rule 13e-100) (Amendment No. 3) Rule 13e-3 Transaction Statement Under Section 13(e) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of the Is

SC 13E3/A SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13E-3 (Rule 13e-100) (Amendment No. 3) Rule 13e-3 Transaction Statement Under Section 13(e) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of the Issuer) Fall Line Endurance Fund, LP (and the affiliated entities and other persons listed on the following page) (Name of Persons Filing Stat

July 13, 2023 SC 14D9/A

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 14D-9 SOLICITATION/RECOMMENDATION STATEMENT UNDER SECTION 14(d)(4) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 3) GreenLight Biosciences Holdings, PBC (Name of

SC 14D9/A #3 UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 14D-9 SOLICITATION/RECOMMENDATION STATEMENT UNDER SECTION 14(d)(4) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 3) GreenLight Biosciences Holdings, PBC (Name of Subject Company) GreenLight Biosciences Holdings, PBC (Name of Person(s) Filing Statement) Common Stock, par value $0.0001 per share (Ti

July 13, 2023 8-K/A

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K/A (Amendment No. 1) CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): May 29, 2023

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K/A (Amendment No. 1) CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): May 29, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction

July 11, 2023 SC 13E3/A

SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13E-3 (Rule 13e-100) (Amendment No. 2) Rule 13e-3 Transaction Statement Under Section 13(e) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of the Is

SC 13E3/A #2 SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13E-3 (Rule 13e-100) (Amendment No. 2) Rule 13e-3 Transaction Statement Under Section 13(e) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of the Issuer) Fall Line Endurance Fund, LP (and the affiliated entities and other persons listed on the following page) (Name of Persons Filing S

July 11, 2023 SC TO-T/A

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE TO TENDER OFFER STATEMENT UNDER SECTION 14(d)(1) OR 13(e)(1) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 1) GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Name of Subje

SC TO-T/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE TO TENDER OFFER STATEMENT UNDER SECTION 14(d)(1) OR 13(e)(1) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 1) GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Name of Subject Company (Issuer)) SW MergerCo, Inc. a wholly-owned subsidiary of (Name of Filing Person — Offeror) SW ParentCo, Inc. an affiliate of (Nam

July 11, 2023 SC 14D9/A

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 14D-9 SOLICITATION/RECOMMENDATION STATEMENT UNDER SECTION 14(d)(4) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 2) GreenLight Biosciences Holdings, PBC (Name of

SC 14D9/A #2 UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 14D-9 SOLICITATION/RECOMMENDATION STATEMENT UNDER SECTION 14(d)(4) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 2) GreenLight Biosciences Holdings, PBC (Name of Subject Company) GreenLight Biosciences Holdings, PBC (Name of Person(s) Filing Statement) Common Stock, par value $0.0001 per share (Ti

June 22, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Lurantos Maria Helena - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G105 (CUSIP Number) Maria H. Lurantos 60 Rantoul Street, #716, Beverly, MA 01915 978-578-7391 (Name, Address and Telephone Number of Person A

June 21, 2023 EX-99.(A)(1)(B)

LETTER OF TRANSMITTAL To accompany certificates of common stock, $0.0001 par value per share, of GreenLight Biosciences Holdings, PBC (“Common Stock”) To Tender Shares of Common Stock of GreenLight Biosciences Holdings, PBC at $0.30 Per Share in Cash

EX-99.(a)(1)(B) Exhibit (a)(1)(B) LETTER OF TRANSMITTAL To accompany certificates of common stock, $0.0001 par value per share, of GreenLight Biosciences Holdings, PBC (“Common Stock”) To Tender Shares of Common Stock of GreenLight Biosciences Holdings, PBC at $0.30 Per Share in Cash Pursuant to the Offer to Purchase dated June 21, 2023 by SW MergerCo, Inc., an entity wholly-owned by SW ParentCo,

June 21, 2023 EX-99.(A)(1)(F)

Notice of Offer to Purchase for Cash All Outstanding Shares of Common Stock GreenLight Biosciences Holdings, PBC $0.30 Net Per Share Pursuant to the Offer to Purchase Dated June 21, 2023 SW MergerCo, Inc., a wholly-owned subsidiary of SW ParentCo, In

EX-99.(a)(1)(F) Exhibit (a)(1)(F) This announcement is neither an offer to purchase nor a solicitation of an offer to sell Shares (as defined below). The Offer (as defined below) is made solely by the Offer to Purchase (as defined below), dated June 21, 2023, and the related Letter of Transmittal (as defined below) and any amendments or supplements thereto. The Offer is not being made to, nor will

June 21, 2023 EX-FILING FEES

Calculation of Filing Fee Tables Schedule TO (Form Type) GreenLight Biosciences Holdings, PBC (Name of Subject Company (Issuer)) SW MergerCo, Inc. (Name of Filing Person — Offeror) a wholly-owned subsidiary of SW ParentCo, Inc. (Name of Filing Person

EX-FILING FEES Exhibit 107 Calculation of Filing Fee Tables Schedule TO (Form Type) GreenLight Biosciences Holdings, PBC (Name of Subject Company (Issuer)) SW MergerCo, Inc.

June 21, 2023 SC TO-T

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE TO TENDER OFFER STATEMENT UNDER SECTION 14(d)(1) OR 13(e)(1) OF THE SECURITIES EXCHANGE ACT OF 1934 GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Name of Subject Company (Issuer

SC TO-T UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE TO TENDER OFFER STATEMENT UNDER SECTION 14(d)(1) OR 13(e)(1) OF THE SECURITIES EXCHANGE ACT OF 1934 GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Name of Subject Company (Issuer)) SW MergerCo, Inc. a wholly-owned subsidiary of (Name of Filing Person — Offeror) SW ParentCo, Inc. an affiliate of (Name of Filing Person —

June 21, 2023 SC 13E3/A

SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13E-3 (Rule 13e-100) (Amendment No. 1) Rule 13e-3 Transaction Statement Under Section 13(e) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of the Is

SC 13E3/A SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13E-3 (Rule 13e-100) (Amendment No. 1) Rule 13e-3 Transaction Statement Under Section 13(e) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of the Issuer) Fall Line Endurance Fund, LP (and the affiliated entities and other persons listed on the following page) (Name of Persons Filing Stat

June 21, 2023 SC 14D9/A

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 14D-9 SOLICITATION/RECOMMENDATION STATEMENT UNDER SECTION 14(d)(4) OF THE SECURITIES EXCHANGE ACT OF 1934 (Amendment No. 1) GreenLight Biosciences Holdings, PBC (Name of

SC 14D9/A Table of Contents UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.

June 21, 2023 EX-99.(A)(1)(E)

Offer to Purchase for Cash All Outstanding Shares of Common Stock GreenLight Biosciences Holdings, PBC $0.30 Net Per Share SW MergerCo, Inc. a wholly-owned subsidiary of SW ParentCo, Inc., an affiliate of Fall Line Endurance Fund, LP

EX-99.(a)(1)(E) Exhibit (a)(1)(E) Offer to Purchase for Cash All Outstanding Shares of Common Stock of GreenLight Biosciences Holdings, PBC at $0.30 Net Per Share by SW MergerCo, Inc. a wholly-owned subsidiary of SW ParentCo, Inc., an affiliate of Fall Line Endurance Fund, LP THE OFFER AND WITHDRAWAL RIGHTS WILL EXPIRE AT ONE MINUTE AFTER 11:59 P.M., NEW YORK CITY TIME, ON JULY 19, 2023, UNLESS TH

June 21, 2023 EX-99.(A)(1)(D)

Offer to Purchase for Cash All Outstanding Shares of Common Stock GreenLight Biosciences Holdings, PBC $0.30 Net Per Share SW MergerCo, Inc. a wholly-owned subsidiary of SW ParentCo, Inc., an affiliate of Fall Line Endurance Fund, LP

EX-99.(a)(1)(D) Exhibit (a)(1)(D) Offer to Purchase for Cash All Outstanding Shares of Common Stock of GreenLight Biosciences Holdings, PBC at $0.30 Net Per Share by SW MergerCo, Inc. a wholly-owned subsidiary of SW ParentCo, Inc., an affiliate of Fall Line Endurance Fund, LP THE OFFER AND WITHDRAWAL RIGHTS WILL EXPIRE AT ONE MINUTE AFTER 11:59 P.M., NEW YORK CITY TIME, ON JULY 19, 2023, UNLESS TH

June 21, 2023 EX-99.(A)(1)(A)

Offer to Purchase for Cash All Outstanding Shares of Common Stock GreenLight Biosciences Holdings, PBC $0.30 Net Per Share SW MergerCo, Inc. A wholly-owned subsidiary of SW ParentCo, Inc., an affiliate of Fall Line Endurance Fund, LP THE OFFER AND WI

EX-99.(a)(1)(A) Table of Contents Exhibit (a)(1)(A) Offer to Purchase for Cash All Outstanding Shares of Common Stock of GreenLight Biosciences Holdings, PBC at $0.30 Net Per Share by SW MergerCo, Inc. A wholly-owned subsidiary of SW ParentCo, Inc., an affiliate of Fall Line Endurance Fund, LP THE OFFER AND WITHDRAWAL RIGHTS WILL EXPIRE AT ONE MINUTE AFTER 11:59 P.M., NEW YORK CITY TIME, ON JULY 1

June 21, 2023 SC 14D9

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 14D-9 SOLICITATION/RECOMMENDATION STATEMENT UNDER SECTION 14(d)(4) OF THE SECURITIES EXCHANGE ACT OF 1934 GreenLight Biosciences Holdings, PBC (Name of Subject Company)

SC 14D9 Table of Contents UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.

June 21, 2023 EX-FILING FEES

Calculation of Filing Fee Tables Schedule 13E-3 (Form Type) GreenLight Biosciences Holdings, PBC (Exact Name of Registrant as Specified in its Charter) Table 1 - Transaction Value Transaction Valuation Fee Rate Amount of Filing Fee Fees to Be Paid $9

EX-FILING FEES Exhibit 107 Calculation of Filing Fee Tables Schedule 13E-3 (Form Type) GreenLight Biosciences Holdings, PBC (Exact Name of Registrant as Specified in its Charter) Table 1 - Transaction Value Transaction Valuation Fee Rate Amount of Filing Fee Fees to Be Paid $9,348,082.

June 21, 2023 SC 13E3

SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13E-3 (Rule 13e-100) Rule 13e-3 Transaction Statement Under Section 13(e) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of the Issuer) Fall Line En

SC 13E3 1 d522502dsc13e3.htm SC 13E3 SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13E-3 (Rule 13e-100) Rule 13e-3 Transaction Statement Under Section 13(e) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of the Issuer) Fall Line Endurance Fund, LP (and the affiliated entities and other persons listed on the following page) (Name of Persons Fi

June 21, 2023 EX-99.(A)(1)(C)

NOTICE OF GUARANTEED DELIVERY For Tender of Shares of Common Stock GreenLight Biosciences Holdings, PBC $0.30 Net Per Share SW MergerCo, Inc. a wholly-owned subsidiary of SW ParentCo, Inc., an affiliate of Fall Line Endurance Fund, LP

EX-99.(a)(1)(C) Exhibit (a)(1)(C) NOTICE OF GUARANTEED DELIVERY For Tender of Shares of Common Stock of GreenLight Biosciences Holdings, PBC at $0.30 Net Per Share by SW MergerCo, Inc. a wholly-owned subsidiary of SW ParentCo, Inc., an affiliate of Fall Line Endurance Fund, LP THE OFFER AND WITHDRAWAL RIGHTS WILL EXPIRE AT ONE MINUTE AFTER 11:59 P.M., NEW YORK CITY TIME, ON JULY 19, 2023, UNLESS T

June 21, 2023 EX-99.(A)(5)(A)

ROTH CAPITAL PARTNERS, LLC 888 SAN CLEMENTE DRIVE, NEWPORT BEACH, CA 92660 | 800.678.9147 | www.roth.com | Member SIPC/FINRA

EX-99.(a)(5)(A) Exhibit (a)(5)(A) May 29, 2023 Special Committee of the Board of Directors GreenLight Biosciences Holdings, PBC 29 Hartwell Avenue Lexington, MA 02421 Members of the Special Committee: You have requested our opinion as to the fairness, from a financial point of view, to holders of common stock, par value $0.0001 per share (“Company Common Stock”), of GreenLight Biosciences Holdings

June 21, 2023 EX-99.(A)(5)(B)

This presentation was prepared by ROTH Capital Partners, LLC (“ROTH MKM”) and provided to the Special Committee of the Board of Directors (the “Special Committee”) of Greenlight Biosciences Holdings, PBC (“GRNA”) in connection with a potential transa

EX-99.(a)(5)(B) May 29, 2023 Discussion Materials for Project Star Wars ROTH CAPITAL PARTNERS, LLC Member FINRA/SIPC www.roth.com Confidential Roth Capital Partners, LLC Corporate Office: 888 San Clemente Drive, Newport Beach, CA 92660 | 949.720.5700 | 800.678.9147 | www.roth.com | Member SIPC / FINRA Regional Offices: Chicago, IL | Los Angeles, CA | Miami Beach, FL | New York, NY | San Francisco,

June 16, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Fall Line Endurance Fund, LP - SC 13D/A Activist Investment

SC 13D/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D/A Under the Securities Exchange Act of 1934 (Amendment No. 3)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Eric O’Brien Fall Line Capital, LLC 160 Bovet Rd., Suite 310, San Mateo, CA 94402 650-235-

June 14, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023

EX-99.1 Exhibit 99.1 EXECUTION VERSION AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS Page ARTICLE I THE MERGER TRANSACTIONS 2 1.1 The Offer 2 1.2 Company Actions 5 1.3 The Merger 7 1.4 Closing 7 1.5 Effective Time 7 1.6 Merger Without Meeting of Stockholders 7 1.7 Effects of the Merg

June 14, 2023 EX-99.2

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.2 Exhibit 99.2 Final Form Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein

June 14, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Furneaux Carol - SC 13D Activist Investment

SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Carol Furneaux 17B Harrison St. Newburyport, MA 01950 978-853-3687 (Name, Address and Telephone

June 13, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Cummings Foundation, Inc. Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) William F. Grant, CFO Cummings Foundation, Inc. 200 W Cummings Park Woburn, MA 01801 781-932-7036 (Name, Address and Tel

June 13, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Anderson Eric Senior Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Eric S. Anderson 25 N Hill Road Westford, MA 01886 781-983-2211 (Name, Address and Telephone Number of Person Authorized

June 13, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Clarke Dennis A. Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Dennis A. Clarke 6 Fernway Terrace Winchester, MA 01890 339-221-8296 (Name, Address and Telephone Number of Person Autho

June 13, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Brewster David B - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G105 (CUSIP Number) David B. Brewster 46 Brewster Street, Cambridge, MA 02138 617-492-2020 (Name, Address and Telephone Number of Person Auth

June 12, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / CG Investments Inc. VI - FORM13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* Greenlight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Larissa Teipner CG Investments Inc. VI 535 Madison Avenue New York City, New York 10022 212-389-8109 (

June 9, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Senczuk Anna Maria - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Anna Senczuk 393 Main St, North Andover, MA 01845 206-910-0636 (Name, Address and Telephone Number of Person Authorized

June 9, 2023 EX-99.3

Joint Filing Agreement pursuant to Rule 13d-1(k)(1) under the Securities Exchange Act of 1934, as amended.

Exhibit 99.3 EXHIBIT A JOINT FILING AGREEMENT The undersigned hereby agree as follows: (i) Each of them is individually eligible to use the Schedule 13D to which this Exhibit is attached, and such Schedule 13D is filed on behalf of each of them; and (ii) Each of them is responsible for the timely filing of such Schedule 13D and any amendments thereto, and for the completeness and accuracy of the i

June 9, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Boscolo Intervest Ltd - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. ) GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Rafael Urquia II, Esq. c/o Fox Horan & Camerini LLP 885 Third Avenue, 17th Floor New York, NY 10022 (21

June 8, 2023 EX-99.3

JOINT FILING AGREEMENT

EX-99.3 EXHIBIT 3 JOINT FILING AGREEMENT In accordance with Rule 13d-1(k) promulgated under the Securities Exchange Act of 1934, as amended, Alfa Holdings, Inc., an British Virgin Islands corporation, and Ricardo A. Sagrera agree to the joint filing on behalf of each of them of a Statement on Schedule 13D (including any amendments thereto) with respect to the common stock, par value $0.0001 per sh

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Steinberg Michael Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Michael Steinberg 5 Azalea Court Scarsdale, NY 10583 (646) 824-1010 (Name, Address and Telephone Number of Person Author

June 8, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023

EX-99.1 2 planofmerger.htm Exhibit 99.1 AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS Page ARTICLE I THE MERGER TRANSACTIONS 2 1.1 The Offer 2 1.2 Company Actions 5 1.3 The Merger 7 1.4 Closing 7 1.5 Effective Time 7 1.6 Merger Without Meeting of Stockholders 7 1.7 Effects of the Mer

June 8, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 NO AGREEMENT, ORAL OR WRITTEN, REGARDING OR RELATING TO ANY OF THE MATTERS COVERED BY THIS DOCUMENT HAS BEEN E

EX-99.1 Exhibit 1 AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 NO AGREEMENT, ORAL OR WRITTEN, REGARDING OR RELATING TO ANY OF THE MATTERS COVERED BY THIS DOCUMENT HAS BEEN ENTERED INTO BETWEEN THE PARTIES. THIS DOCUMENT IS INTENDED SOLELY TO FACILITATE DISCUSSIONS AMONG THE PARTIES IDENTIFIED HEREIN.

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Gillian Jason Michael - SEC 13D - GREENLIGHT AND JASON M. GILLIAN Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Jason M. Gillian 50 Beverly Dr Durham NC, 27707 919 538 4340 (Name, Address and Telephone Number of Person Authorized to

June 8, 2023 EX-1

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Zahir Fares - EXHIBIT 1

Exhibit 1 JOINT FILING AGREEMENT In accordance with Rule 13d-1(k)(1) promulgated under the Securities Exchange Act of 1934, as amended, the undersigned hereby agree that they are jointly filing this statement on Schedule 13D.

June 8, 2023 EX-99.3

JOINT FILING AGREEMENT

EX-99.3 Exhibit 99.3 JOINT FILING AGREEMENT The undersigned hereby agree that they are filing this statement jointly pursuant to Rule 13d-1(k)(1). Each of them is responsible for the timely filing of such Schedule 13D and any amendments thereto, and for the completeness and accuracy of the information concerning such person contained therein; but none of them is responsible for the completeness or

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / ALEXANDRIA REAL ESTATE EQUITIES, INC. - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Dean A. Shigenaga Chief Financial Officer Alexandria Venture Investments, LLC 26 North Euclid Avenue P

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / S2G Advisors, LLC - SC 13D Activist Investment

SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Information to be included in statements filed pursuant to Rule 13d-1(a) and amendments thereto filed pursuant to Rule 13d-2(a) Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / DHAMANKAR HIMANSHU HEMANT - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Himanshu Hemant Dhamankar c/o GreenLight Biosciences Holdings, PBC 29 Hartwell Avenue Lexington, MA 02421 (Name, Address

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / RUETTGERS MICHAEL C - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Michael Ruettgers 453 Bedford Road Carlisle, MA 01741 978-369-4886 (Name, Address and Telephone Number

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Zahir Fares - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G105 (CUSIP Number) Fares Zahir c/o TMF Nominees Ltd, 2nd Floor, The Grand Pavilion, Commercial Centre, 802 West Bay Road,

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Sagar Rosemary Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Rosemary Sagar 1643 Brickell Ave Miami, FL 33129 (646) 420-5715 (Name, Address and Telephone Number of Person Authorized

June 8, 2023 EX-99.4

JOINT FILING AGREEMENT

EX-99.4 EXHIBIT 4 JOINT FILING AGREEMENT In accordance with Rule 13d-1(k) promulgated under the Securities Exchange Act of 1934, as amended, Alfa Holdings, Inc., an British Virgin Islands corporation, and Ricardo A. Sagrera agree to the joint filing on behalf of each of them of a Statement on Schedule 13D (including any amendments thereto) with respect to the common stock, par value $0.0001 per sh

June 8, 2023 EX-99.3

JOINT FILING AGREEMENT

EX-99.3 Exhibit 99.3 JOINT FILING AGREEMENT The undersigned hereby agree that they are filing this statement jointly pursuant to Rule 13d-1(k)(1). Each of them is responsible for the timely filing of such Schedule 13D and any amendments thereto, and for the completeness and accuracy of the information concerning such person contained therein; but none of them is responsible for the completeness or

June 8, 2023 EX-99.2

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.2 Exhibit 2 Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [ ] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein shall have t

June 8, 2023 EX-99.2

SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT

Exhibit 99.2 SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT This Secured Convertible Note Purchase Agreement, dated as of May 29, 2023 (this “Agreement”), is entered into by and among SW ParentCo, Inc., a Delaware corporation (the “Company”), and the undersigned persons and entities (each an “Investor” and, collectively, the “Investors”). WHEREAS, the Company has entered into that c

June 8, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS

Exhibit 99.1 AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS Page ARTICLE I THE MERGER TRANSACTIONS 2 1.1 The Offer 2 1.2 Company Actions 5 1.3 The Merger 7 1.4 Closing 7 1.5 Effective Time 7 1.6 Merger Without Meeting of Stockholders 7 1.7 Effects of the Merger 7 1.8 Certificate of In

June 8, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023

EX-99.1 Exhibit 99.1 EXECUTION VERSION AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS Page ARTICLE I THE MERGER TRANSACTIONS 2 1.1 The Offer 2 1.2 Company Actions 5 1.3 The Merger 7 1.4 Closing 7 1.5 Effective Time 7 1.6 Merger Without Meeting of Stockholders 7 1.7 Effects of the Merg

June 8, 2023 EX-99.3

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.3 4 contribexchangeagreement.htm Exhibit 99.3 Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [ ] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used b

June 8, 2023 EX-99.4

Joint Filing Agreement

Exhibit 99.4 Joint Filing Agreement In accordance with Rule 13d-1(k) promulgated under the Securities Exchange Act of 1934, as amended, the undersigned hereby agree to the joint filing with all other Reporting Persons (as such term is defined in the Schedule 13D referred to below) on behalf of each of them of a statement on Schedule 13D (including amendments thereto) with respect to the Common Sto

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Gonzalez May Carlos Alfredo - SC 13D Activist Investment

SC 13D 1 d377652dsc13d.htm SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. N/A)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Carlos A. Gonzalez May c/o Rivas Capital LLC 104 Mt Auburn St, Su

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Iwuchukwu Ifeyinwa Jane - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Ifeyinwa Iwuchukwu 9 Wyman Road, Billerica MA 01821 865-621-6760 (Name, Address and Telephone Number of Person Authorize

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Macadino Caitlin - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Caitlin Macadino 8 Copley Place, Billerica, MA 01821 617-543-7717 (Name, Address and Telephone Number of Person Authoriz

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Serum Life Sciences Ltd - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Serum Life Sciences Ltd 15 Grosvenor Street London England W1K 4QZ (Name, Address and Telephone Number of Person Authori

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Sagrera Ricardo A. - SC 13D Activist Investment

SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. N/A)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Ricardo A. Sagrera 801 Brickell Avenue Miami, Florida 33131 (786) 252-7993 (Name, Address an

June 8, 2023 EX-99.3

AGREEMENT REGARDING THE JOINT FILING OF SCHEDULE 13D

Exhibit 99.3 AGREEMENT REGARDING THE JOINT FILING OF SCHEDULE 13D Pursuant to Rule 13(d)-1(k)(1) promulgated under the Securities Exchange Act of 1934, as amended, each of the undersigned acknowledges and agrees that the foregoing statement on Schedule 13D is filed on behalf of the undersigned and that all subsequent amendments to this statement on Schedule 13D shall be filed on behalf of the unde

June 8, 2023 EX-99.1

Joint Filing Agreement (filed herewith).

Exhibit 99.1 JOINT FILING AGREEMENT The undersigned acknowledge and agree that the foregoing statement on Schedule 13D is filed on behalf of each of the undersigned and that all subsequent amendments to this statement on Schedule 13D may be filed on behalf of each of the undersigned without the necessity of filing additional joint filing agreements. The undersigned acknowledge that each shall be r

June 8, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023

EX-99.1 Exhibit 99.1 EXECUTION VERSION AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS Page ARTICLE I THE MERGER TRANSACTIONS 2 1.1 The Offer 2 1.2 Company Actions 5 1.3 The Merger 7 1.4 Closing 7 1.5 Effective Time 7 1.6 Merger Without Meeting of Stockholders 7 1.7 Effects of the Merg

June 8, 2023 EX-99.2

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023

EX-99.2 Exhibit 99.2 AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 NO AGREEMENT, ORAL OR WRITTEN, REGARDING OR RELATING TO ANY OF THE MATTERS COVERED BY THIS DOCUMENT HAS BEEN ENTERED INTO BETWEEN THE PARTIES. THIS DOCUMENT IS INTENDED SOLELY TO FACILITATE DISCUSSIONS AMONG THE PARTIES IDENTIFIED HERE

June 8, 2023 EX-99.3

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.3 Exhibit 99.3 Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [ ] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein shall hav

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Cormorant Global Healthcare Master Fund, LP Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock (Title of Class of Securities) 39536G105 (CUSIP Number) Neb Obradovic Cormorant Asset Management, LP 200 Clarendon Street, 52nd Floor Boston, MA 02116 Tel. No.: 857-702-0386 (Name, Address

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Gupta Sweta Kumari - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Sweta Gupta 102 Harlow St, Arlington 02474 617-306-3375 (Name, Address and Telephone Number of Person Authorized to Rece

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / RAMACHANDRIYA DHARMAN KARTHIKEYAN - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Karthikeyan Ramachandriya Dharman 8 Cardinal Street, Winchester, MA 01890 405-338-8660 (Name, Address and Telephone Numb

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Skizim Nicholas John Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Nicholas J. Skizim 22 Crowley Ave, Dedham, MA 02026 713-201-2961 (Name, Address and Telephone Number of Person Authorize

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / RiverRoad Capital Partners, LLC - SC 13D Activist Investment

SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Gregory T. Lucier RiverRoad Capital Partners, LLC 510 E. Hyman Avenue, Suite 21 Aspen, CO 81611 (760) 481-5664 Wi

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Board of Trustees of The Leland Stanford Junior University - SC 13D Activist Investment

SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Michael Taylor The Board of Trustees of the Leland Stanford Junior University 635 Knight Way St

June 8, 2023 EX-99.2

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.2 3 d462932dex992.htm EX-99.2 Exhibit 99.2 Final Form Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms

June 8, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 NO AGREEMENT, ORAL OR WRITTEN, REGARDING OR RELATING TO ANY OF THE MATTERS COVERED BY THIS DOCUMENT HAS BEEN E

EX-99.1 Exhibit 1 AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 NO AGREEMENT, ORAL OR WRITTEN, REGARDING OR RELATING TO ANY OF THE MATTERS COVERED BY THIS DOCUMENT HAS BEEN ENTERED INTO BETWEEN THE PARTIES. THIS DOCUMENT IS INTENDED SOLELY TO FACILITATE DISCUSSIONS AMONG THE PARTIES IDENTIFIED HEREIN.

June 8, 2023 EX-99.2

SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT

Exhibit 99.2 SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT This Secured Convertible Note Purchase Agreement, dated as of May 29, 2023 (this “Agreement”), is entered into by and among SW ParentCo, Inc., a Delaware corporation (the “Company”), and the undersigned persons and entities (each an “Investor” and, collectively, the “Investors”). WHEREAS, the Company has entered into that c

June 8, 2023 EX-99.3

CONTRIBUTION AND EXCHANGE AGREEMENT

Exhibit 99.3 Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [ ] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein shall have the me

June 8, 2023 EX-99.3

CONTRIBUTION AND EXCHANGE AGREEMENT

Exhibit 99.3 Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [ ] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein shall have the me

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Aulisa Lorenzo - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Lorenzo Aulisa 9 Whitetail Way, Bangor, ME 04401 314-546-6444 (Name, Address and Telephone Number of Person Authorized t

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / CONTINENTAL GRAIN CO - SCHEDULE 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Continental Grain Company 767 Fifth Avenue, 15th Floor New York, NY 10153-0015 212-207-5100 Attention:

June 8, 2023 EX-99.2

CONTRIBUTION AND EXCHANGE AGREEMENT

Exhibit 99.2 Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of May 29, 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and Conti Greenlight Investors, LP (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not d

June 8, 2023 EX-99.2

SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT

EX-99.2 Exhibit 2 SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT This Secured Convertible Note Purchase Agreement, dated as of May 29, 2023 (this “Agreement”), is entered into by and among SW ParentCo, Inc., a Delaware corporation (the “Company”), and the undersigned persons and entities (each an “Investor” and, collectively, the “Investors”). WHEREAS, the Company has entered into t

June 8, 2023 EX-99.2

SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT

Exhibit 99.2 SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT This Secured Convertible Note Purchase Agreement, dated as of May 29, 2023 (this “Agreement”), is entered into by and among SW ParentCo, Inc., a Delaware corporation (the “Company”), and the undersigned persons and entities (each an “Investor” and, collectively, the “Investors”). WHEREAS, the Company has entered into that c

June 8, 2023 EX-99.2

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.2 Exhibit 99.2 Final Form Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Insud Pharma S.L. - SCHEDULE 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* Greenlight Biosciences Holdings, PBC (Name of Issuer) Common stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) c/o Ana Mondedeu Insunza Insud Pharma, S.L. Calle de Manuel Pombo Angulo, No. 28, 3rd Floor 28050 Madr

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Prelude Ventures LLC - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Tim Woodward One Ferry Building, Suite 300 San Francisco, CA 94111 415-729-1270 (Name, Address and Tel

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Tao Capital Management LP - SC 13D Activist Investment

SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Lori D. Mills Tao Capital Management LP 1 Letterman Drive, Suite C4-420 San Francisco, CA 94129

June 8, 2023 SC 13D

ENVI / Environmental Impact Acquisition Corp - Class A / Senczuk Anna Maria - SCHEDULE 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Anna Senczuk 393 Main St, North Andover, MA 01845 206-910-0636 (Name, Address and Telephone Number of Person Authorized

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / New Stuff, LLC - SC 13D Activist Investment

SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Theodore J. Theophilos Two North Riverside Plaza, Suite 1240 Chicago, Illinois 60606 (312) 466-

June 8, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Lewis & Clark Ventures I, LP - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Meg Lane 120 S. Central Avenue, Suite 1000 St. Louis, MO 63105 (314) 392-5259 (Name, Address and Telephone Number of Per

June 8, 2023 EX-99.3

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.3 Exhibit 3 Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [ ] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein shall have t

June 8, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023

Exhibit 99.1 AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS Page ARTICLE I THE MERGER TRANSACTIONS 2 1.1 The Offer 2 1.2 Company Actions 5 1.3 The Merger 7 1.4 Closing 7 1.5 Effective Time 7 1.6 Merger Without Meeting of Stockholders 7 1.7 Effects of the Merger 7 1.8 Certificate of In

June 8, 2023 EX-99.1

JOINT FILING AGREEMENT

EX-99.1 Exhibit 99.1 JOINT FILING AGREEMENT In accordance with Rule 13d-1(k)(1) promulgated under the Securities Exchange Act of 1934, as amended, the undersigned agree to the joint filing on behalf of each of them of a Schedule 13D (including any and all amendments thereto) with respect to the common stock, par value $0.0001 per share, of GreenLight Biosciences Holdings, PBC, and further agree th

June 8, 2023 EX-99.1

CONTRIBUTION AND EXCHANGE AGREEMENT

Exhibit 99.1 Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of May 29, 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and Continental Grain Company (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not define

June 7, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Richard Roger David - SCHEDULE 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Roger Richard 118 Stallion Hill Rd, Fiskdale, MA 01518 774-757-7109 (Name, Address and Telephone Number of Person Author

June 7, 2023 EX-99.1

Joint Filing Agreement

EX-99.1 2 d502744dex991.htm EX-99.1 Exhibit 99.1 CUSIP No. 39536G 105 Page 11 of 11 Joint Filing Agreement In accordance with Rule 13d-1(k) of the Securities Exchange Act of 1934, as amended, the undersigned agree to the joint filing on behalf of each of them of a Statement on Schedule 13D (including any and all amendments thereto) with respect to the common stock, par value $0.0001 per share (the

June 7, 2023 EX-99.2

SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT

EX-99.2 Exhibit 99.2 SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT This Secured Convertible Note Purchase Agreement, dated as of May 29, 2023 (this “Agreement”), is entered into by and among SW ParentCo, Inc., a Delaware corporation (the “Company”), and the undersigned persons and entities (each an “Investor” and, collectively, the “Investors”). WHEREAS, the Company has entered int

June 7, 2023 EX-99.3

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.3 Table of Contents Exhibit 99.3 Confidential Draft Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of May 29, 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and Morningside Venture Investments Ltd. (the “Rollover Investor”), on the other hand. Except as otherwise

June 7, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023

Exhibit 99.1 AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS Page ARTICLE I THE MERGER TRANSACTIONS 2 1.1 The Offer 2 1.2 Company Actions 5 1.3 The Merger 7 1.4 Closing 7 1.5 Effective Time 7 1.6 Merger Without Meeting of Stockholders 7 1.7 Effects of the Merger 7 1.8 Certificate of In

June 7, 2023 EX-99.1

JOINT FILING AGREEMENT

EX-99.1 CUSIP No. 39536G105 EXHIBIT 99.1 JOINT FILING AGREEMENT The undersigned, being duly authorized thereunder, hereby execute this agreement as an exhibit to this Schedule 13D amendment to evidence the agreement of the below-named parties, in accordance with the rules promulgated pursuant to the Securities Exchange Act of 1934, to file this Schedule 13D amendment (including amendments thereto)

June 7, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Morningside Venture Investments Ltd - SC 13D/A Activist Investment

SC 13D/A CUSIP No. 39536G105 UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. 1)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G105 (CUSIP Number) Morningside Venture Investments Limited C/O THC Management Services S.A.M

June 7, 2023 EX-99.2

SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT

Exhibit 99.2 SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT This Secured Convertible Note Purchase Agreement, dated as of May 29, 2023 (this “Agreement”), is entered into by and among SW ParentCo, Inc., a Delaware corporation (the “Company”), and the undersigned persons and entities (each an “Investor” and, collectively, the “Investors”). WHEREAS, the Company has entered into that c

June 7, 2023 EX-99.3

CONTRIBUTION AND EXCHANGE AGREEMENT

Exhibit 99.3 Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [ ] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein shall have the me

June 7, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Jeremy & Hannelore Grantham Environmental Trust - SC 13D Activist Investment

SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Ramsay Ravenel The Jeremy and Hannelore Grantham Environmental Trust 53 State Street, 33rd Floo

June 7, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Series Greenlight, a separate series of BlueIO Growth LLC Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Jan Haas 200 Summit Drive, 2nd Floor Burlington, Massachusetts 01803 (617) 899-0294 (Name, Address and Telephone Number

June 6, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Ortega-Valle Marta - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Marta Ortega-Valle c/o GreenLight Biosciences Holdings, PBC 29 Hartwell Avenue Lexington, MA 02421 (617) 616-8188 (Name,

June 6, 2023 EX-99.3

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.3 Exhibit 99.3 Final Form Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein

June 6, 2023 EX-99.1

Joint Filing Agreement

EX-99.1 2 d467456dex991.htm EX-99.1 Exhibit 99.1 Joint Filing Agreement The undersigned hereby agree that a single Schedule 13D (or any amendment thereto) relating to the Common Stock of GreenLight Biosciences Holdings, PBC shall be filed on behalf of each of the undersigned and that this Agreement shall be filed as an exhibit to such Schedule 13D. IN WITNESS WHEREOF, the undersigned hereby execut

June 6, 2023 EX-99.2

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023

EX-99.2 3 d467456dex992.htm EX-99.2 Exhibit 99.2 EXECUTION VERSION AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS Page ARTICLE I THE MERGER TRANSACTIONS 2 1.1 The Offer 2 1.2 Company Actions 5 1.3 The Merger 7 1.4 Closing 7 1.5 Effective Time 7 1.6 Merger Without Meeting of Stockholde

June 6, 2023 SC 13D

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / MLS Capital Fund II LP - SC 13D Activist Investment

SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Gregory Young MLSCF Management (Labuan), LLP Suite 26.03, Level 26, GTower 199, Jalan Tun Razak

June 5, 2023 EX-99.2

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.2 Exhibit 99.2 Final Form Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein

June 5, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023

EX-99.1 Exhibit 99.1 EXECUTION VERSION AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS Page ARTICLE I THE MERGER TRANSACTIONS 2 1.1 The Offer 2 1.2 Company Actions 5 1.3 The Merger 7 1.4 Closing 7 1.5 Effective Time 7 1.6 Merger Without Meeting of Stockholders 7 1.7 Effects of the Merg

June 5, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / KODIAK VENTURE PARTNERS III LP - SC 13D/A Activist Investment

SC 13D/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D/A Under the Securities Exchange Act of 1934 (Amendment No. 1)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) David Furneaux Kodiak Ventures Management Company, Inc. 11 Peter Grover Rd Bethel, ME 0421

May 31, 2023 8-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): May 30, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporation) (

May 30, 2023 EX-2.1

Agreement and Plan of Merger, dated May 29, 2023, by and among Issuer, Parent and Merger Sub (incorporated by reference to Exhibit 2.1 to the GreenLight Biosciences Holdings, PBC Form 8-K filed with the Securities and Exchange Commission on May 30, 2023)

EX-2.1 Exhibit 2.1 AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 TABLE OF CONTENTS Page ARTICLE I THE MERGER TRANSACTIONS 2 1.1 The Offer 2 1.2 Company Actions 5 1.3 The Merger 7 1.4 Closing 7 1.5 Effective Time 7 1.6 Merger Without Meeting of Stockholders 7 1.7 Effects of the Merger 7 1.8 Certificate

May 30, 2023 8-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): May 29, 2023 GreenLight Bioscienc

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): May 29, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporation) (

May 30, 2023 EX-99.1

AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 NO AGREEMENT, ORAL OR WRITTEN, REGARDING OR RELATING TO ANY OF THE MATTERS COVERED BY THIS DOCUMENT HAS BEEN E

EX-99.1 2 d474103dex991.htm EX-99.1 Exhibit 99.1 AGREEMENT AND PLAN OF MERGER by and among SW PARENTCO, INC. SW MERGERCO, INC. and GreenLight Biosciences Holdings, PBC Dated as of May 29, 2023 NO AGREEMENT, ORAL OR WRITTEN, REGARDING OR RELATING TO ANY OF THE MATTERS COVERED BY THIS DOCUMENT HAS BEEN ENTERED INTO BETWEEN THE PARTIES. THIS DOCUMENT IS INTENDED SOLELY TO FACILITATE DISCUSSIONS AMONG

May 30, 2023 EX-99.3

CONTRIBUTION AND EXCHANGE AGREEMENT

EX-99.3 Exhibit 99.3 Privileged & Confidential CONTRIBUTION AND EXCHANGE AGREEMENT THIS CONTRIBUTION AND EXCHANGE AGREEMENT (this “Agreement”) is made as of [], 2023, by and between SW ParentCo, Inc., a Delaware corporation (“Parent”), on the one hand, and [ ] (the “Rollover Investor”), on the other hand. Except as otherwise set forth herein, capitalized terms used but not defined herein shall hav

May 30, 2023 EX-99.2

SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT

EX-99.2 Exhibit 99.2 SW PARENTCO, INC. SECURED CONVERTIBLE NOTE PURCHASE AGREEMENT This Secured Convertible Note Purchase Agreement, dated as of May 29, 2023 (this “Agreement”), is entered into by and among SW ParentCo, Inc., a Delaware corporation (the “Company”), and the undersigned persons and entities (each an “Investor” and, collectively, the “Investors”). WHEREAS, the Company has entered int

May 30, 2023 SC14D9C

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 14D-9 (Rule 14d-101) Solicitation/Recommendation Statement Under Section 14(d)(4) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Su

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 14D-9 (Rule 14d-101) Solicitation/Recommendation Statement Under Section 14(d)(4) of the Securities Exchange Act of 1934 GreenLight Biosciences Holdings, PBC (Name of Subject Company) GreenLight Biosciences Holdings, PBC (Name of Persons Filing Statement) Common Stock, Par Value $0.0001 Per Share (Title of Class of Se

May 30, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Fall Line Endurance Fund, LP - SC 13D/A Activist Investment

SC 13D/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D/A Under the Securities Exchange Act of 1934 (Amendment No. 2)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Eric O’Brien Fall Line Capital, LLC 119 South B Street, San Mateo, CA 94401 650-520-6790 (

May 30, 2023 EX-99.1

GreenLight Biosciences Enters into Definitive Merger Agreement with Consortium Led by Fall Line Endurance Fund for Go-Private Transaction

EX-99.1 Exhibit 99.1 GreenLight Biosciences Enters into Definitive Merger Agreement with Consortium Led by Fall Line Endurance Fund for Go-Private Transaction Lexington, Mass., May 30, 2023 — GreenLight Biosciences Holdings, PBC (NASDAQ: GRNA) (“GreenLight” or the “Company”) today announced that the Company has entered into a definitive agreement (the “Merger Agreement”) to be acquired by a group

May 30, 2023 EX-4.1

Form of Advance Note

EX-4.1 Exhibit 4.1 THE SECURITY REPRESENTED HEREBY HAS NOT BEEN REGISTERED UNDER THE SECURITIES ACT OF 1933, AS AMENDED, OR APPLICABLE STATE SECURITIES LAWS, AND MAY NOT BE OFFERED OR SOLD IN THE ABSENCE OF AN EFFECTIVE REGISTRATION STATEMENT UNDER SUCH FEDERAL AND STATE SECURITIES LAWS OR AN EXEMPTION FROM REGISTRATION THEREUNDER. THIS NOTE IS SUBJECT TO (I) THAT CERTAIN SUBORDINATION AGREEMENT D

May 11, 2023 10-Q

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 10-Q

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 10-Q (Mark One) ☒ QUARTERLY REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the quarterly period ended March 31, 2023 OR ☐ TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the transition period from to Commission File Number: 001-39894 GreenLight Biosciences Holdings, PBC (Exact Name of Registrant as Specified in its Charter) Delaware 85-1914700 (State or other jurisdiction of incorporation or organization) (I.

May 11, 2023 EX-99

GreenLight Biosciences Announces First Quarter 2023 Financial Results and Highlights Recent Company Progress

EX-99 2 grna-ex991.htm EX-99.1 GreenLight Biosciences Announces First Quarter 2023 Financial Results and Highlights Recent Company Progress BOSTON, May 11, 2023—GreenLight Biosciences, PBC (Nasdaq: GRNA), a biotechnology company dedicated to making RNA products affordable and accessible for human health and agriculture, today reported operational highlights and financial results for the first quar

May 11, 2023 8-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): May 11, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporation) (

March 30, 2023 EX-99

GreenLight Biosciences Holdings Confirms Receipt of Indication of Interest from Fall Line

GreenLight Biosciences Holdings Confirms Receipt of Indication of Interest from Fall Line BOSTON, Mass.

March 30, 2023 SC 13D/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Fall Line Endurance Fund, LP - SC 13D/A Activist Investment

SC 13D/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D/A Under the Securities Exchange Act of 1934 (Amendment No. 1)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G 105 (CUSIP Number) Eric O’Brien Fall Line Capital, LLC 119 South B Street, San Mateo, CA 94401 650-520-6790 (

March 30, 2023 8-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): March 30, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporation)

March 28, 2023 EX-10

Employment Offer Letter, dated February 7, 2021, between GreenLight Biosciences, Inc. and Amin Khan.

Certain identified information has been omitted from this exhibit because it is not material and of the type that the reigstrant treats as private or confidential.

March 28, 2023 EX-21

List of Subsidiaries

Exhibit 21.1 GreenLight Biosciences Holdings, PBC List of Subsidiaries Name of Subsidiary* Jurisdiction GreenLight Biosciences, Inc. (100%) Delaware, United States of America GreenLight Security Corporation (100%) Massachusetts, United States of America GreenLight Biosciences Espana, S.L. (100%) Spain * Percentage in parentheses indicates GreenLight Bioscience Holdings, PBC’s direct or indirect pe

March 28, 2023 POS AM

As filed with the Securities and Exchange Commission on March 28, 2023

POS AM Table of Contents As filed with the Securities and Exchange Commission on March 28, 2023 Registration Statement No.

March 28, 2023 EX-10

Lease Agreement, dated March 9, 2022 between ARE-MA Region No.8, LLC GreenLight Biosciences, Inc.

DocuSign Envelope ID: 3D9421BF-E23B-4F57-8C3D-FC854D1C296D LEASE AGREEMENT  DocuSign Envelope ID: 3D9421BF-E23B-4F57-8C3D-FC854D1C296D DocuSign Envelope ID: 3D9421BF-E23B-4F57-8C3D-FC854D1C296D THIS LEASE AGREEMENT (this “Lease”) is made this 9day of March, 2022, between ARE-MA REGION NO.

March 28, 2023 8-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): March 28, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporation)

March 28, 2023 EX-99

GreenLight Biosciences Announces Fourth Quarter and Full Year 2022 Financial Results

GreenLight Biosciences Announces Fourth Quarter and Full Year 2022 Financial Results - Two agriculture solutions now under regulatory review, pending EPA approval: Calantha™, a solution to protect potatoes, and a separate solution to protect honeybees from Varroa mite - Updated clinical strategy to accelerate development of its next-generation Covid vaccine candidate; decision to advance universal

March 28, 2023 EX-10

Collaboration and License Agreement, by and between the Company and EpiVax Therapeutics, Inc., dated January 8, 2023

Execution Version Certain identified information has been omitted from this exhibit because it is not material and of the type that the reigstrant treats as private or confidential.

March 28, 2023 10-K

Form 10-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 10-K (Mark One) ☒ ANNUAL REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the fiscal year ended December 31, 2022 OR ☐ TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 FOR THE TRANSITION PERIOD FROM TO Commission File Number 001-39894 GreenLight Bios

March 9, 2023 8-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): March 09, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporation)

March 9, 2023 EX-99

GreenLight Biosciences Outlines Development Strategy and Highlights Portfolio Updates at Human Health R&D Day -- Progressing human health pipeline conferring mRNA platform advantages for infectious disease and personalized oncology medicines; pursuin

GreenLight Biosciences Outlines Development Strategy and Highlights Portfolio Updates at Human Health R&D Day - Progressing human health pipeline conferring mRNA platform advantages for infectious disease and personalized oncology medicines; pursuing capital efficient strategy targeting unmet medical needs, in both the developed and lower- and middle-income countries (LMICs), pairing focused R&D spend with partnerships - Accelerating development of next-generation COVID vaccine candidate; decision to advance universal pan-sarbecovirus vaccine candidate in place of mono-valent Wuhan based antigen.

March 9, 2023 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-267619 Prospectus Supplement No. 6 (To Prospectus dated October 5, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 6 (this “Prospectus Supplement”) amends and supplements the prospectus dated October 5, 2022 (as amen

March 7, 2023 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-267619 Prospectus Supplement No. 5 (To Prospectus dated October 5, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 5 (this “Prospectus Supplement”) amends and supplements the prospectus dated October 5, 2022 (as amen

March 7, 2023 8-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): March 07, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporation)

March 7, 2023 EX-99

GreenLight Biosciences Outlines Development Strategy and Highlights Portfolio Updates at Plant Health R&D Day

EX-99 2 grna-ex991.htm EX-99.1 GreenLight Biosciences Outlines Development Strategy and Highlights Portfolio Updates at Plant Health R&D Day BOSTON, March 7, 2023 — GreenLight Biosciences (Nasdaq: GRNA), a public benefit corporation striving to deliver on the full potential of RNA to address some of the world’s toughest problems in human health and agriculture, will today outline its pipeline stra

February 14, 2023 SC 13G/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / Cormorant Asset Management, LP Passive Investment

SC 13G/A 1 schedule13g.htm UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13G Under the Securities Exchange Act of 1934 (Amendment No. 1)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock (Title of Class of Securities) 39536G105 (CUSIP Number) December 31, 2022 (Date of Event which Requires Filing of this Statement) Check the appropriate box to d

February 14, 2023 SC 13G/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / S2G Advisors, LLC - SC 13G/A Passive Investment

SC 13G/A UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13G Under the Securities Exchange Act of 1934 (Amendment No. 1)* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G105 (CUSIP Number) 12/31/2022 (Date of Event Which Requires Filing of this Statement) Check the appropriate box

February 1, 2023 8-K

Regulation FD Disclosure, Financial Statements and Exhibits, Other Events

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): January 27, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporatio

February 1, 2023 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-267619 Prospectus Supplement No. 4 (To Prospectus dated October 5, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 4 (this “Prospectus Supplement”) amends and supplements the prospectus dated October 5, 2022 (as amen

February 1, 2023 EX-99.1

GreenLight Biosciences receives approval to initiate Phase I/II clinical trial of Covid-19 mRNA vaccine candidate

GreenLight Biosciences receives approval to initiate Phase I/II clinical trial of Covid-19 mRNA vaccine candidate • The Rwanda FDA has given regulatory approval to GreenLight to start a Phase I/II clinical trial of its Covid-19 vaccine candidate in Rwanda • This phase I/II clinical trial is the first step in GreenLight’s integrated strategy of bringing a universal Covid-19 mRNA vaccine to market globally.

January 30, 2023 SC 13G/A

US39536G1058 / GREENLIGHT BIOSCIENCES HOLDINGS PBC A / HIGHBRIDGE CAPITAL MANAGEMENT LLC - GREENLIGHT BIOSCIENCES HOLDINGS, PBC Passive Investment

SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13G/A Under the Securities Exchange Act of 1934 (Amendment No. 2)* GreenLight Biosciences Holdings, PBC (formerly known as Environmental Impact Acquisition Corp.) (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G105 (CUSIP Number) December 31, 2022 (Date of event which requires filing

January 9, 2023 8-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): January 08, 2023 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporatio

January 9, 2023 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-267619 Prospectus Supplement No. 3 (To Prospectus dated October 5, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 3 (this “Prospectus Supplement”) amends and supplements the prospectus dated October 5, 2022 (as amen

January 9, 2023 EX-99.01

GreenLight Biosciences and EpiVax Therapeutics Sign Exclusive Collaboration Agreement to Develop Personalized Cancer Vaccines

GreenLight Biosciences and EpiVax Therapeutics Sign Exclusive Collaboration Agreement to Develop Personalized Cancer Vaccines ● The effort will combine GreenLight’s mRNA design and manufacturing expertise with EpiVax’s advanced Ancer® pipeline for personalized cancer vaccine design ● EpiVax Therapeutics, Inc.

November 9, 2022 10-Q

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 10-Q

c UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 10-Q (Mark One) ? QUARTERLY REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the quarterly period ended September 30, 2022 OR ? TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the transition period from to Commission File Number: 001-39894 GreenLight Biosciences Holdings, PBC (Exact Name of Registrant as Specified in its Charter) Delaware 85-1914700 (State or other jurisdiction of incorporation or organization) (I.

November 9, 2022 8-K

Financial Statements and Exhibits, Results of Operations and Financial Condition

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): November 09, 2022 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporati

November 9, 2022 EX-99.PR

GREENLIGHT BIOSCIENCES HOLDINGS, PBC Condensed Consolidated Statements of Operations (unaudited) (In thousands, except share and per share amounts) GREENLIGHT BIOSCIENCES HOLDINGS, PBC Condensed Consolidated Balance Sheets Data (unaudited) (In thousa

GreenLight Biosciences Announces Third Quarter 2022 Financial Results and Highlights Recent Company Progress ? Working toward clinical trial initiation for COVID vaccine candidate in 2023 ? Shingles vaccine candidate selection planned for 2023 in partnership with Serum Institute of India ? Calantha?, Colorado potato beetle solution, commercial launch planned for 2023, subject to regulatory approval ? Honeybee solution EPA submission planned for 2023 ? Strategic realignment focused on near-term value drivers.

November 9, 2022 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-267619 Prospectus Supplement No. 2 (To Prospectus dated October 5, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 2 (this ?Prospectus Supplement?) updates, amends and supplements the prospectus dated October 5, 2022

October 12, 2022 8-K

Regulation FD Disclosure, Financial Statements and Exhibits, Costs Associated with Exit or Disposal Activities

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): October 11, 2022 GreenLight Biosciences Holdings, PBC (Exact name of Registrant as Specified in Its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporatio

October 12, 2022 EX-99.PRESSRELEASE

GreenLight Biosciences announces realignment to focus on near-term value drivers

GreenLight Biosciences announces realignment to focus on near-term value drivers BOSTON, MA, October 12, 2022?GreenLight?Biosciences (Nasdaq: GRNA), a public benefit corporation striving to bring effective and safe solutions to make food clean and affordable for everyone and dedicated to developing health solutions for every person on our planet, today announced a realignment to focus on key near-term value drivers and extend its cash runway.

October 12, 2022 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-267619 Prospectus Supplement No. 1 (To Prospectus dated October 5, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 1 (this ?Prospectus Supplement?) amends and supplements the prospectus dated October 5, 2022 (as amen

October 6, 2022 424B3

GreenLight Biosciences Holdings, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Table of Contents Filed Pursuant to Rule 424(b)(3) Registration No. 333-267619 PROSPECTUS GreenLight Biosciences Holdings, PBC 114,692,259 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus relates to the resale, from time to time, of up to 114,692,259 shares of our common stock, par value $0.0001 per share (?Common Stock?), by the selling s

October 4, 2022 CORRESP

GreenLight Biosciences Holdings, PBC 200 Boston Avenue Medford, Massachusetts 02155

GreenLight Biosciences Holdings, PBC 200 Boston Avenue Medford, Massachusetts 02155 October 4, 2022 VIA EDGAR Securities and Exchange Commission Division of Corporation Finance 100 F Street, N.

October 3, 2022 LETTER

LETTER

United States securities and exchange commission logo October 3, 2022 Andrey Zarur President and Chief Executive Officer GreenLight Biosciences Holdings, PBC 200 Boston Avenue Medford, MA 02155 Re: GreenLight Biosciences Holdings, PBC Registration Statement on Form S-1 Filed September 26, 2022 File No.

September 26, 2022 EX-10.35

First Amendment to Commercial Lease, dated September 12, 2022, by and between ARE-NC REGION NO. 17, LLC and GreenLight Biosciences Inc. (incorporated by reference to Exhibit 10.35 to the Registration Statement on Form S-1 filed with the Securities and Exchange Commission on September 26, 2022).

Exhibit 10.35 FIRST AMENDMENT TO LEASE THIS FIRST AMENDMENT TO LEASE (this ?First Amendment?) is made as of September 12, 2022, by and between ARE-NC REGION NO. 17, LLC, a Delaware limited liability company (?Landlord?), and GREENLIGHT BIOSCIENCES INC., a Delaware corporation (?Tenant?). RECITALS A. Landlord and Tenant are now parties to that certain Lease Agreement dated as of September 30, 2021,

September 26, 2022 EX-FILING FEES

Filing Fee Table

Exhibit 107 Calculation of Filing Fee Tables Form S-1 (Form Type) GreenLight Biosciences Holdings, PBC (Exact Name of Registrant as Specified in its Charter) Table 1: Newly Registered and Carry Forward Securities Security Type Security Class Title Fee Calculation or Carry Forward Rule Amount Registered(1)(2) Proposed Maximum Offering Price Per Unit Maximum Aggregate Offering Price Fee Rate Amount of Registration Fee Newly Registered Securities Fees to Be Paid Equity Common stock, par value $0.

September 26, 2022 EX-10.34

Commercial Lease, dated March 9, 2022, by and between ARE-MA Region No. 8, LLC and GreenLight Biosciences Inc. (incorporated by reference to Exhibit 10.34 to the Registration Statement on Form S-1 filed with the Securities and Exchange Commission on September 26, 2022).

Exhibit 10.34 LEASE AGREEMENT THIS LEASE AGREEMENT (this ?Lease?) is made this 9 day of March, 2022, between ARE-MA REGION NO. 8, LLC, a Delaware limited liability company (?Landlord?), and GREENLIGHT BIOSCIENCES INC., a Delaware corporation (?Tenant?). Building: 29 Hartwell Avenue, Lexington, Massachusetts Premises: The entire Building, containing approximately 59,000 rentable square feet, as det

September 26, 2022 EX-99.2

MANAGEMENT’S DISCUSSION AND ANALYSIS OF FINANCIAL CONDITION AND RESULTS OF OPERATIONS OF GREENLIGHT

Exhibit 99.2 MANAGEMENT?S DISCUSSION AND ANALYSIS OF FINANCIAL CONDITION AND RESULTS OF OPERATIONS OF GREENLIGHT The following discussion and analysis of the financial condition and results of operations of GreenLight Biosciences, Inc. and its consolidated subsidiaries should be read together with GreenLight?s audited consolidated financial statements as of and for the years ended December 31, 202

September 26, 2022 8-K

Financial Statements and Exhibits, Other Events

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): September 26, 2022 GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Exact name of registrant as specified in its charter) Delaware 001-39894 85-1914700 (State or other jurisdiction of incorporat

September 26, 2022 EX-99.1

INDEX TO FINANCIAL STATEMENTS Page Financial Statements for GreenLight Biosciences, Inc. (accounting predecessor to GreenLight Biosciences Holdings, PBC): Report of Independent Registered Public Accounting Firm F-2 Consolidated Balance Sheets as of D

Exhibit 99.1 INDEX TO FINANCIAL STATEMENTS Page Financial Statements for GreenLight Biosciences, Inc. (accounting predecessor to GreenLight Biosciences Holdings, PBC): Report of Independent Registered Public Accounting Firm F-2 Consolidated Balance Sheets as of December 31, 2021 and 2020 F-3 Consolidated Statements of Operations for the years ended December 31, 2021 and 2020 F-4 Consolidated State

September 26, 2022 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-262574 Prospectus Supplement No. 4 (To Prospectus dated April 6, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 4 (this ?Prospectus Supplement?) updates, amends and supplements the prospectus dated April 6, 2022 (as

September 26, 2022 S-1

As filed with the Securities and Exchange Commission on September 2 6 , 2022 Registration No. 333- UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM S-1 REGISTRATION STATEMENT UNDER THE SECURITIES ACT OF 1933 GreenLight Bio

Table of Contents As filed with the Securities and Exchange Commission on September 2 6 , 2022 Registration No.

September 26, 2022 EX-10.36

Third Amendment to Commercial Lease, dated September 12, 2022, by and between ARE-NC REGION NO. 17, LLC and GreenLight Biosciences Inc. (incorporated by reference to Exhibit 10.36 to the Registration Statement on Form S-1 filed with the Securities and Exchange Commission on September 26, 2022).

Exhibit 10.36 THIRD AMENDMENT TO LEASE THIS THIRD AMENDMENT TO LEASE (this ?Third Amendment?) is made as of September 12, 2022, by and between ARE-NC REGION NO. 17, LLC, a Delaware limited liability company (?Landlord?), and GREENLIGHT BIOSCIENCES INC., a Delaware corporation (?Tenant?). RECITALS A. Landlord and Tenant are now parties to that certain Lease Agreement dated as of January 15, 2019, a

August 15, 2022 EX-10.1

Form of Securities Subscription Agreement, dated as of August 11, 2022, between GreenLight Biosciences Holdings, PBC and each purchaser named in the signature pages thereto (incorporated by reference to Exhibit 10.1 to the Current Report on Form 8-K filed with the Securities and Exchange Commission on August 15, 2022).

Exhibit 10.1 SUBSCRIPTION AGREEMENT August 11, 2022 GreenLight Biosciences Holdings, PBC, a Delaware corporation (formerly known as Environmental Impact Acquisition Corp.) (the ?Company?), is seeking commitments from interested investors to purchase shares of the Company?s Common Stock, par value $0.0001 per share (the ?Common Stock?), for a purchase price of $[]1 per share (the ?Purchase Price pe

August 15, 2022 EX-99.1

GreenLight Biosciences Announces Second Quarter 2022 Financial Results and Highlights Recent Company Progress

Exhibit 99.1 GreenLight Biosciences Announces Second Quarter 2022 Financial Results and Highlights Recent Company Progress ? GreenLight raised $108.4 million in private placement led by S2G Ventures with strong participation from existing and new investors with cash expected to fund critical programs through the first half of 2023. ? GreenLight and Samsung Biologics completed their first commercia

August 15, 2022 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-262574 Prospectus Supplement No. 3 (To Prospectus dated April 6, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 3 (this ?Prospectus Supplement?) updates, amends and supplements the prospectus dated April 6, 2022 (as

August 15, 2022 10-Q

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 10-Q

c UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 10-Q (Mark One) ? QUARTERLY REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the quarterly period ended June 30, 2022 OR ? TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the transition period from to Commission File Number: 001-39894 GreenLight Biosciences Holdings, PBC (Exact Name of Registrant as Specified in its Charter) Delaware 85-1914700 (State or other jurisdiction of incorporation or organization) (I.

August 15, 2022 8-K

Entry into a Material Definitive Agreement, Unregistered Sales of Equity Securities, Financial Statements and Exhibits

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): August 11, 2022 GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Exact Name of Registrant as Specified in its Charter) Delaware 001-39894 85-1914700 (State or Other Jurisdiction of Incorporation) (Commission File Number) (IRS Employer Identification No.

August 1, 2022 8-K

Regulation FD Disclosure, Financial Statements and Exhibits

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): August 1, 2022 GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Exact name of registrant as specified in its charter) Delaware 001-39894 85-1914700 (State or other jurisdiction of incorporation)

August 1, 2022 EX-99.1

GreenLight Biosciences and Samsung Biologics complete first commercial-scale engineering run for mRNA Covid-19 vaccine

Exhibit 99.1 GreenLight Biosciences and Samsung Biologics complete first commercial-scale engineering run for mRNA Covid-19 vaccine ? GreenLight?s messenger RNA production process is transferable to large-scale equipment and CMO facilities ? Technology transfer and scale-up from lab bench to Samsung?s commercial facility was completed in seven months ? GreenLight?s mRNA synthesis reaction had a ti

July 29, 2022 8-K

Regulation FD Disclosure, Financial Statements and Exhibits

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): July 29, 2022 GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Exact name of registrant as specified in its charter) Delaware 001-39894 85-1914700 (State or other jurisdiction of incorporation)

July 29, 2022 EX-99.1

NIH, GreenLight Biosciences to collaborate on COVID-19 vaccine development for new variants

Exhibit 99.1 NIH, GreenLight Biosciences to collaborate on COVID-19 vaccine development for new variants BOSTON, July 29, 2022?GreenLight Biosciences has announced a collaboration with the National Institutes of Health (NIH) to develop COVID-19 vaccines that are more broadly protective against new variants and with longer-lasting effects. ?This is a tremendous opportunity for us to partner with a

June 1, 2022 10-Q/A

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 10-Q/A (Amendment No. 1)

c UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 10-Q/A (Amendment No.

June 1, 2022 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-262574 Prospectus Supplement No. 2 (To Prospectus dated April 6, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 2 (this ?Prospectus Supplement?) updates, amends and supplements the prospectus dated April 6, 2022 (as

May 18, 2022 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-262574 Prospectus Supplement No. 1 (To Prospectus dated April 6, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 1 (this ?Prospectus Supplement?) updates, amends and supplements the prospectus dated April 6, 2022 (as

May 16, 2022 10-Q

UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 10-Q

c UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, DC 20549 FORM 10-Q (Mark One) ? QUARTERLY REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the quarterly period ended March 31, 2022 OR ? TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the transition period from to Commission File Number: 001-39894 GreenLight Biosciences Holdings, PBC (Exact Name of Registrant as Specified in its Charter) Delaware 85-1914700 (State or other jurisdiction of incorporation or organization) (I.

April 12, 2022 EX-99.1

GreenLight Biosciences Holdings, PBC 2022 Equity and Incentive Plan

EXHIBIT 99.1 GREENLIGHT BIOSCIENCES HOLDINGS, PBC 2022 EQUITY AND INCENTIVE PLAN Section 1. Purposes of the Plan The purposes of the GreenLight Biosciences Holdings, PBC 2022 Equity and Incentive Plan (the ?Plan?) are to (i) provide long-term incentives and rewards to those employees, officers, directors and other key persons (including consultants) of GreenLight Biosciences Holdings, PBC (the ?Co

April 12, 2022 S-8

As filed with the Securities and Exchange Commission on April 12, 2022

As filed with the Securities and Exchange Commission on April 12, 2022 Registration No.

April 12, 2022 EX-99.2

GreenLight Biosciences Holdings, PBC 2022 Employee Stock Purchase Plan (incorporated by reference to Exhibit 99.2 to the Registration Statement on Form S-8 filed with the Securities and Exchange Commission on April 12, 2022).

EXHIBIT 99.2 GREENLIGHT BIOSCIENCES HOLDINGS, PBC 2022 EMPLOYEE STOCK PURCHASE PLAN 1. Purpose of the Plan The purpose of the GreenLight Biosciences Holdings, PBC 2022 Employee Stock Purchase Plan (the ?Plan?) is to provide eligible employees of GreenLight Biosciences Holdings, PBC (the ?Company?) and each Designated Company (as defined below) with the opportunity to purchase shares of the Company

April 12, 2022 EX-FILING FEES

Filing Fee Table

Exhibit 107 Calculation of Filing Fee Tables Form S-8 (Form Type) GreenLight Biosciences Holdings, PBC (Exact Name of Registrant as Specified in its Charter) Table 1: Newly Registered Securities Security Type Security Class Title Fee Calculation Rule Amount Registered(1) Proposed Maximum Offering Price Per Share Maximum Aggregate Offering Price Fee Rate Amount of Registration Fee Equity Common stock, par value $0.

April 7, 2022 424B3

GreenLight Biosciences Holdings, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Table of Contents Filed Pursuant to Rule 424(b)(3) Registration No. 333-262574 PROSPECTUS GreenLight Biosciences Holdings, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus relates to the resale, from time to time, of up to 86,631,958 shares of our common stock, par value $0.0001 per share (?Common Stock?), by the selling sec

April 5, 2022 EX-10.32

License Agreement, dated March 10, 2022, between GreenLight Biosciences, Inc. and Serum Institute of India Private Limited (incorporated by reference to Exhibit 10.32 to Post-Effective Amendment No. 1 to Form S-1 Registration Statement filed with the Securities and Exchange Commission on April 5, 2022).

Exhibit 10.32 Confidential Certain identified information has been omitted from this exhibit because it is not material and of the type that the registrant treats as private or confidential. [***] indicates that information has been omitted. LICENSE AGREEMENT BETWEEN SERUM INSTITUTE OF INDIA PRIVATE LIMITED AND GREENLIGHT BIOSCIENCES, INC. Confidential LICENSE AGREEMENT This License Agreement (thi

April 5, 2022 POS AM

As filed with the Securities and Exchange Commission on April 5, 2022 Registration No. 333-262574 UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 POST-EFFECTIVE AMENDMENT NO. 1 TO FORM S-1 REGISTRATION STATEMENT UNDER THE SECU

Table of Contents As filed with the Securities and Exchange Commission on April 5, 2022 Registration No.

March 31, 2022 EX-99.1

INDEX TO FINANCIAL STATEMENTS Page Financial Statements for GreenLight Biosciences, Inc. (accounting predecessor to GreenLight Biosciences Holdings, PBC): Report of Independent Registered Public Accounting Firm F-2 Consolidated Balance Sheets as of D

Exhibit 99.1 INDEX TO FINANCIAL STATEMENTS Page Financial Statements for GreenLight Biosciences, Inc. (accounting predecessor to GreenLight Biosciences Holdings, PBC): Report of Independent Registered Public Accounting Firm F-2 Consolidated Balance Sheets as of December 31, 2021 and 2020 F-3 Consolidated Statements of Operations for the years ended December 31, 2021 and 2020 F-4 Consolidated State

March 31, 2022 EX-99.2

MANAGEMENT’S DISCUSSION AND ANALYSIS OF FINANCIAL CONDITION AND RESULTS OF OPERATIONS OF GREENLIGHT

Exhibit 99.2 MANAGEMENT?S DISCUSSION AND ANALYSIS OF FINANCIAL CONDITION AND RESULTS OF OPERATIONS OF GREENLIGHT The following discussion and analysis of the financial condition and results of operations of GreenLight Biosciences, Inc. and its consolidated subsidiaries should be read together with GreenLight?s audited consolidated financial statements as of and for the years ended December 31, 202

March 31, 2022 10-K

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 10-K (Mark One) ☒ ANNUAL REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the fiscal year ended December 31, 2021 OR ☐ TRANSITION REPORT PUR

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 10-K (Mark One) ? ANNUAL REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the fiscal year ended December 31, 2021 OR ? TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934 For the transition period from to Commission file number 001-39894 GREENLIGHT BIOS

March 31, 2022 8-K/A

Financial Statements and Exhibits

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 8-K/A (Amendment No. 1) CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): February 2, 2022 GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Exact name of registrant as specified in its charter) Delaware 001-39894 85-1914700 (State or other jurisdic

March 31, 2022 EX-4.3

Description of the Company’s Securities Registered Pursuant to Section 12 of the Securities Exchange Act of 1934 (incorporated by reference to Exhibit 4.3 to the Registration Statement on Form 10-K filed with the Securities and Exchange Commission on March 31, 2022).

Exhibit 4.3 DESCRIPTION OF THE REGISTRANT?S SECURITIES REGISTERED PURSUANT TO SECTION 12 OF THE SECURITIES EXCHANGE ACT OF 1934 The following description summarizes certain important terms of the securities of GreenLight Biosciences Holdings, PBC, formerly Environmental Impact Acquisition Corp. (?we,? ?us,? ?our,? the ?Company? or ?New GreenLight?) as of March 15, 2022. Because the following descr

March 17, 2022 424B3

GREENLIGHT BIOSCIENCES HOLDINGS, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Filed Pursuant to Rule 424(b)(3) Registration No. 333-262574 Prospectus Supplement No. 1 (To Prospectus dated February 14, 2022) GREENLIGHT BIOSCIENCES HOLDINGS, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus supplement no. 1 (this ?Prospectus Supplement?) updates, amends and supplements the prospectus dated February 14, 2

March 17, 2022 8-K

Entry into a Material Definitive Agreement

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): March 11, 2022 GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Exact name of registrant as specified in its charter) Delaware 001-39894 85-1914700 (State or other jurisdiction of incorporation)

February 25, 2022 SC 13D

GRNA / GreenLight Biosciences Inc / Fall Line Endurance GP, LLC Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, DC 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No.

February 15, 2022 424B3

GreenLight Biosciences Holdings, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants

Table of Contents Filed Pursuant to Rule 424(b)(3) Registration No. 333-262574 PROSPECTUS GreenLight Biosciences Holdings, PBC 86,631,958 Shares of Common Stock 10,350,000 Shares of Common Stock Issuable Upon Exercise of Warrants This prospectus relates to the resale, from time to time, of up to 86,631,958 shares of our common stock, par value $0.0001 per share (?Common Stock?), by the selling sec

February 15, 2022 424B3

424B3

February 14, 2022 EX-99.6

GREENLIGHT BIOSCIENCES, INC. ENVIRONMENTAL IMPACT ACQUISITION CORP.

Exhibit 99.6 GREENLIGHT BIOSCIENCES, INC. ENVIRONMENTAL IMPACT ACQUISITION CORP. December 21, 2021 MVIL, LLC c/o 22nd Floor, Hang Lung Centre 2-20 Paterson St., Causeway Bay, HK Re: Payment of Purchase Price under Subscription Agreement Ladies and Gentlemen: This letter will confirm our agreement that pursuant to and effective as of your purchase of one or more Convertible Instruments (the ?Instru

February 14, 2022 EX-99.1

JOINT FILING AGREEMENT

Exhibit 99.1 JOINT FILING AGREEMENT This Joint Filing Agreement, dated as of February 14, 2022, is by and among Cormorant Global Healthcare Master Fund, LP, Cormorant Global Healthcare GP, LLC, Cormorant Private Healthcare Fund II, LP, Cormorant Private Healthcare GP II, LLC, Cormorant Asset Management, LP and Bihua Chen (collectively, the ?Filers?). Each of the Filers may be required to file with

February 14, 2022 EX-99.5

CONVERTIBLE INVESTMENT INSTRUMENT AGREEMENT

Exhibit 99.5 CONVERTIBLE INVESTMENT INSTRUMENT AGREEMENT This Convertible Investment Instrument Agreement (?Agreement?) is dated as of December 21, 2021 by and among GreenLight Biosciences, Inc., a Delaware corporation (the ?Company?) and the individuals and entities who become parties to this Agreement by executing and delivering an Agreement Signature Page in the form of Exhibit A hereto in acco

February 14, 2022 EX-99.1

JOINT FILING AGREEMENT

CUSIP No. 39536G105 EXHIBIT 99.1 JOINT FILING AGREEMENT The undersigned, being duly authorized thereunder, hereby execute this agreement as an exhibit to this Schedule 13D to evidence the agreement of the below-named parties, in accordance with the rules promulgated pursuant to the Securities Exchange Act of 1934, to file this Schedule 13D (including amendments thereto) jointly on behalf of each s

February 14, 2022 SC 13D

GRNA / GreenLight Biosciences Inc / Morningside Venture Investments Ltd - SC 13D Activist Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G105 (CUSIP Number) Morningside Venture Investments Limited C/O THC Management Services S.A.M., 2nd Floor, Le Prince De Gal

February 14, 2022 SC 13G

ENVIRONMENTAL IMPACT ACQUISITION CORP / FIR TREE CAPITAL MANAGEMENT LP Passive Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, DC 20549 SCHEDULE 13G Under the Securities Exchange Act of 1934 (Amendment No.

February 14, 2022 SC 13G

GRNA / GreenLight Biosciences Inc / Cormorant Asset Management, LP Passive Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13G Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock (Title of Class of Securities) 39536G105 (CUSIP Number) February 2, 2022 (Date of Event which Requires Filing of this Statement) Check the appropriate box to designate the rule pursuant to

February 11, 2022 EX-99.A

Joint Filing Agreement

Exhibit A Joint Filing Agreement The undersigned hereby agree that a single Schedule 13D (or any amendment thereto) relating to the Common Stock of GreenLight Biosciences Holdings, PBC shall be filed on behalf of each of the undersigned and that this Agreement shall be filed as an exhibit to such Schedule 13D.

February 11, 2022 SC 13D

GRNA / GreenLight Biosciences Inc / KODIAK VENTURE PARTNERS III LP - SC 13D Activist Investment

SC 13D UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13D Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock (Title of Class of Securities) 39536G 105 (CUSIP Number) (Name, Address and Telephone Number of Person Authorized to Receive Notices and Communications) February 2, 2022 (Date of Ev

February 11, 2022 SC 13G

GRNA / GreenLight Biosciences Inc / S2G Advisors, LLC - SC 13G Passive Investment

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13G Under the Securities Exchange Act of 1934 (Amendment No. )* GreenLight Biosciences Holdings, PBC (Name of Issuer) Common Stock, par value $0.0001 per share (Title of Class of Securities) 39536G105 (CUSIP Number) 02/02/2022 (Date of Event Which Requires Filing of this Statement) Check the appropriate box to designa

February 10, 2022 SC 13G/A

ENVIRONMENTAL IMPACT ACQUISITION CORP / ADAGE CAPITAL PARTNERS GP, L.L.C. - ENVIRONMENTAL IMPACT ACQUISITION CORP. Passive Investment

SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13G/A Under the Securities Exchange Act of 1934 (Amendment No. 1)* Environmental Impact Acquisition Corp. (Name of Issuer) Class A Common Stock, $0.0001 par value per share (Title of Class of Securities) 29408N106 (CUSIP Number) December 31, 2021 (Date of Event Which Requires Filing of This Statement) Check the appropriate box to d

February 10, 2022 CORRESP

GreenLight Biosciences Holdings, PBC 200 Boston Avenue Medford, Massachusetts 02155

GreenLight Biosciences Holdings, PBC 200 Boston Avenue Medford, Massachusetts 02155 February 10, 2022 VIA EDGAR Securities and Exchange Commission Division of Corporation Finance 100 F Street, N.

February 9, 2022 SC 13G/A

ENVIRONMENTAL IMPACT ACQUISITION CORP / HIGHBRIDGE CAPITAL MANAGEMENT LLC - ENVIRONMENTAL IMPACT ACQUISITION CORP. Passive Investment

SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 SCHEDULE 13G/A Under the Securities Exchange Act of 1934 (Amendment No. 1)* Environmental Impact Acquisition Corp. (Name of Issuer) Class A Common Stock, par value $0.0001 per share (Title of Class of Securities) 29408N106 (CUSIP Number) December 31, 2021 (Date of event which requires filing of this statement) Check the appropriate box to d

February 9, 2022 LETTER

LETTER

United States securities and exchange commission logo February 9, 2022 Andrey Zarur President and Chief Executive Officer GreenLight Biosciences Holdings, PBC 200 Boston Avenue Medford, Massachusetts 02155 Re: GreenLight Biosciences Holdings, PBC Registration Statement on Form S-1 Filed February 7, 2022 File No.

February 7, 2022 S-1

As filed with the Securities and Exchange Commission on February 7, 2022 Registration No. 333- UNITED STATES SECURITIES AND EXCHANGE COMMISSION WASHINGTON, D.C. 20549 FORM S-1 REGISTRATION STATEMENT UNDER THE SECURITIES ACT OF 1933 GreenLight Bioscie

Table of Contents As filed with the Securities and Exchange Commission on February 7, 2022 Registration No.

February 7, 2022 8-K

Regulation FD Disclosure, Changes in Control of Registrant, Departure of Directors or Certain Officers; Election of Directors; Appointment of Certain Officers; Compensatory Arrangements of Certain Officers, Amendments to Articles of Incorporation or Bylaws; Change in Fiscal Year, Amendment to Registrant's Code of Ethics, or Waiver of a Provision of the Code of Ethics, Change in Shell Company Status, Entry into a Material Definitive Agreement, Material Modification to Rights of Security Holders, Unregistered Sales of Equity Securities, Financial Statements and Exhibits, Changes in Registrant's Certifying Accountant, Creation of a Direct Financial Obligation or an Obligation under an Off-Balance Sheet Arrangement of a Registrant, Completion of Acquisition or Disposition of Assets

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 FORM 8-K CURRENT REPORT Pursuant to Section 13 or 15(d) of the Securities Exchange Act of 1934 Date of Report (Date of earliest event reported): February 2, 2022 GREENLIGHT BIOSCIENCES HOLDINGS, PBC (Exact name of registrant as specified in its charter) Delaware 001-39894 85-1914700 (State or other jurisdiction of incorporatio

February 7, 2022 EX-16.1

Letter from WithumSmith+Brown, PC to the U.S. Securities and Exchange Commission.

Exhibit 16.1 February 7, 2022 Office of the Chief Accountant Securities and Exchange Commission 100 F Street, NE Washington, D.C. 20549 Ladies and Gentlemen: We have read GreenLight Biosciences Holding, PBC?s (formerly known as Environmental Impact Acquisition Corp.) statements included under Item 4.01 of its Form 8-K dated February 2, 2022. We agree with the statements concerning our Firm under I

February 7, 2022 EX-3.1

Certificate of Incorporation of GreenLight Biosciences Holdings, PBC.

Exhibit 3.1 SECOND AMENDED AND RESTATED CERTIFICATE OF INCORPORATION OF ENVIRONMENTAL IMPACT ACQUISITION CORP. February 2, 2022 Environmental Impact Acquisition Corp., a corporation organized and existing under the laws of the State of Delaware (the ?Corporation?), DOES HEREBY CERTIFY AS FOLLOWS: 1. The name of the Corporation is ?Environmental Impact Acquisition Corp.?. The original certificate o

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